ON THE COVER

Wang et al. (pages 1373–1387) show that cytokinin signaling activates de novo WUSCHEL expression in the Arabidopsis leaf axil, enabling axillary meristem initiation. This requires a permissive epigenetic environment and is directly activated by B-type ARABIDOPSIS RESPONSE REGULATOR transcription factors. The cover image shows localization of the cytokinin signaling reporter TCS (green) and WUSCHEL (red) in a transverse cross section of an Arabidopsis shoot apex.