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COVER: Organogenesis of nitrogen-fixing nodules in legume plants is initiated in specific root cortical cells in response to Rhizobium infection. In this issue, Complainville et al. (pages 2778-2791) explore cell-to-cell communication processes that occur during nodule initiation in Medicago subspecies. In uninoculated Medicago plants, plasmodesmata-mediated communication between root phloem and cortical cells is restricted. The authors show that nodule initiation induces symplasmic continuity between the phloem and nodule initials that is regulated throughout nodule development and may be a critical element in the control of nodule organogenesis. The cover image shows a transverse section of a nodule primordium from a Rhizobium-inoculated Medicago truncatula plant expressing green fluorescent protein under the control of a phloem-specific promoter. Strong green fluorescent protein fluorescence is visible not only in the phloem but in all cell types involved in the the formation of the nodule primordium.
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