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Blumeria graminis is an obligate biotrophic fungus that infects barley and other cereals and causes a powdery mildew disease. Both et al. (pages 2107-2122) conducted gene expression profiling of primary metabolism genes in B. graminis f sp hordei throughout the in planta infection cycle in barley and report that there are striking patterns of coordinate expression among numerous fungal genes in defined metabolic pathways. The results allowed the authors to assess the metabolic status of the fungus as it infects the host plant and completes its life cycle. The cover image shows barley epidermal cells infected with B. graminis. Fungal haustoria, structures used by the fungus to absorb nutrients from the host, are visible as green fluorescent structures labeled with a WGA-Alexa 488 conjugate. The outline of the host cell walls are autofluorescent blue.